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The original AU's plot is very similar to Undertale's. All of the characters, (Except humans, which are rodents (ex. Chara is a rat and Frisk is a mouse)) are " Felines ", various species of cats.
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The AU Feline!Tale itself is more of a Multi-verse, containing smaller "Sub AUS" inside it (ex. Under!Fell, Under!Swap) however to know is a Sub-AU is in Feline!Tale or not, you are to add an !FT at the end of it.(ex. Under!Fell!FT, or Under!Swap!FT). A Sub-AU can be as small as a tiny town, or as big as an entire Undertale map.
There are Non-Adoptive and Adoptive felines in this AU. Adoptive AUS are AUS that come from the world called Shiziku.(). They fly on a plane that travels into Feline!Tale. Most often pick "roles", or characters from Non Feline!Tale AUS they want to be before getting there. They often study on the "role" before going there.
Adoptive AUS are AUS that were born into Feline!Tale in a Sub-AU. They stay in that Sub-AU until taught or stumble across how to cross the barrier into another Sub-AU. All Adoptive AUS are born with ear tufts, however many cut them off, as for they are unable to grow back.
Travelling from Sub-AU to Sub-AU is not uncommon. Many AUs gather to one Sub-AU and attend school there. It is not uncommon to have, for example, a Fell!Feline meet an Ink!Feline at a school. Glitches are not uncommon either, for many of them are Non-Adoptive Characters that accidentally had some sort of accident while flying into Feline!Tale.